The Prowler can be used to improve anaerobic abilities by performing sprints that are short in duration and high in intensity. It can also aid aerobic conditioning by performing reps longer in duration or reducing rest between sets.
The prowler is also of benefit in improving strength, power, and speed. This can be achieved by adding weight (as much as possible) and explosively performing pushing/pulling drills. This is of obvious benefit in sports and movements where you are required to perform explosive and dynamic movements (sprinting, jumping, bounding, planting, rapid changing of direction).
Furthermore, the prowler can help establish proper motor patterns and strengthens muscles that are critical for running (the muscles in the feet and lower legs, quads, hamstrings, glutes, and core). It can also dramatically improve your ability to accelerate in all directions.